Madison Jeffries possesses psionic powers that can affect metal, plastic, and glass. He can psionically levitate objects made of any of these three substances. Jeffries can psionically rearrange parts of machinery into various forms, and he is particularly skilled at psionically restructuring machinery into duplicating the form and functions of parts of the human body. Jeffries can also psionically manipulate metal on the atomic and molecular levels.

When the Amazing X-Men attempted to assist in the Sentinel Evacuation of Northern America, Box reprogrammed the Sentinels to target the X-Men, though the X-Men were later able to reverse Box's efforts. Box was able to conceal himself and pose as a human during the evacuation along with Copycat. They were discovered at a checkpoint and provided a distraction for a group of Altered Humans to pass through, but were subsequently hunted down by Weapon X and Jean Grey with the former slaying Box in the process.